Cow Cave is an important Quaternary paleontological site in Chudleigh Gorge, Devon, UK., now known to have a sequence of cave-earths and stalagmite floors that range in age from Upper Middle Pleistocene (~MIS 7 interglacial) through to the Holocene (Flandrian) and the present day. Excavations in 1927-1934, and again in 1962-3, have provided a rich fauna, with some artefacts. Here, the stratigraphy of the deposits is now more fully described and the faunal remains are considered in their stratigraphical contexts. Data supporting the existence of former cave entrances are presented along with an analysis of the processes of sedimentation and taphonomy with their climatic interpretations. Based on recent U-Th dating of a critical Stalagmite horizon, a chronology of the mid-Pleistocene to Holocene sequence is discussed. Finally, further excavation in Cow Cave and nearby sites is recommended.