The invertebrate faunas of four caves in northern Venezuela were studied. Three were dry and one was a wet cave; but guano provided the base of the food chain in all four caves. The faunas in each were strikingly different. Of 24 non-accidental species of arthropods, all were judged to be troglophiles. All were scavengers except for 5 predators, one ectoparasite, and one parasitoid.
Peck, Stewart B..
A contribution to the knowledge of the invertebrate cave faunas of Venezuela: Invertebrate faunas of tropical American caves, part 4.
International Journal of Speleology,
Available at: http://scholarcommons.usf.edu/ijs/vol12/iss1/8